A mysterious vehicle using a giant magnet steals a highly potent mutagen known as PX-41 from a secret laboratory in the Arctic Circle. The Anti-Villain League (AVL), led by Silas Ramsbottom, sends one of his agents, Lucy Wilde to collect Gru, who was former super-villain. Meanwhile, at Gru's house, it is one of his daughters Agnes's birthday and hosts a party for her, Later Lucy Wilde uses her lipstick taser to get Gru to the AVL headquarters, two minions witness the kidnapping and try to save Gru but get electrocuted by Lucy, When they make it at the AVL Silas Ramsbottom introduces himself (much to Gru and the minions amusement) and tells him he was taken there to track down the culprit of the person who stole the lab and recover the mutagen. However, Gru refuses, claiming that he is now a legitimate businessman of making jelly out of every fruit there is but the jelly turns out to taste horrible and the devoted father to Margo and Edith. Longing to resume his criminal life, Dr. Nefario, Gru's friend and assistant, leaves Gru for new employment. Gru reluctantly agrees to investigate the theft and works with undercover AVL agent Lucy Wilde.
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